Fin Whale
From Whalepedia.org
The Fin Whale (Balaenoptera physalus), also called the Finback Whale or Razorback or Common Rorqual, is a marine mammal belonging to the suborder of baleen whales. It is the second largest whale and the second largest living animal after the Blue Whale,[2] growing to nearly 27 meters (88 ft) long.[2]
Long and slender, the Fin Whale's body is brownish-grey with a paler underside. There are at least two distinct subspecies: the Northern Fin Whale of the North Atlantic, and the larger Antarctic Fin Whale of the Southern Ocean. It is found in all the world's major oceans, from polar to tropical waters. It is absent only from waters close to the ice pack at both the north and south poles and relatively small areas of water away from the open ocean. The highest population density occurs in temperate and cool waters.[3] Its food consists of small schooling fish, squid and crustaceans including mysids and krill.
Like all other large whales, the Fin Whale was heavily hunted during the twentieth century and is an endangered species. The International Whaling Commission (IWC) has issued a moratorium on commercial hunting of this whale,[4] although Iceland and Japan have announced intentions to resume hunting, the latter country stating it will kill a quota of 50 whales for the 2008 season. Collisions with ships and noise from human activity are also significant threats to the recovery of the species.
Contents |
Taxonomy
Fin Whales are rorquals (family Balaenopteridae), a family that includes the Humpback Whale, the Blue Whale, the Bryde's Whale, the Sei Whale and the Minke Whale. The family Balaenopteridae diverged from the other families of the suborder Mysticeti as long ago as the middle Miocene.[6] However, it is not known when the members of these families diverged from each other. Hybridization between the Blue Whale and the Fin Whale is known to occur at least occasionally in the North Atlantic[7] and in the North Pacific.[8]
As of 2006, there are two named subspecies, each with distinct physical features and vocalizations. B. p. physalus (Linnaeus 1758), or Northern Fin Whale, is found in the North Atlantic, and B. p. quoyi (Fischer 1829), or Antarctic Fin Whale, is found in the Southern Ocean.[9] Most experts consider the Fin Whales of the North Pacific to be a third unnamed subspecies.[3] On a global scale, the three groups rarely mix, if at all.
Description and behaviour
The whale has a series of 56–100 pleats or grooves along the bottom of the body that run from the tip of the chin to the navel that allow the throat area to expand greatly during feeding. It has a curved, prominent (60 cm, 24 in) dorsal fin about three-quarters of the way along the back. Its flippers are small and tapered, and its tail is wide, pointed at the tip, and notched in the centre.[2]
When the whale surfaces, the dorsal fin is visible soon after the spout. The spout is vertical and narrow and can reach heights of 6 metres.[11] The whale will blow one to several times on each visit to the surface, staying close to the surface for about one and a half minutes each time. The tail remains submerged during the surfacing sequence. It then dives to depths of up to 250 metres (820 ft), each dive lasting between 10 and 15 minutes. Fin Whales have been known to leap completely out of the water.[11]
Life history
Mating occurs in temperate, low-latitude seas during the winter, and the gestation period is eleven months to one year. A newborn weans from its mother at 6 or 7 months of age when it is 11 or 12 metres (36 to 39 ft) in length, and the calf follows the mother to the winter feeding ground. Females reproduce every 2 to 3 years, with as many as 6 foetuses being reported, but single births are far more common. Females reach sexual maturity at between 3 and 12 years of age.[11]
Feeding
Behaviour
The Fin Whale is one of the fastest cetaceans and can sustain speeds of 37 kilometres per hour (23 mph, 20 knots),[11] and bursts in excess of 40 kilometres per hour (25 mph, 22 knots) have been recorded, earning the Fin Whale the nickname "the greyhound of the deep".[14] Fin Whales are more gregarious than other rorquals, and often live in groups of 6–10 individuals, although on the feeding grounds aggregations of up to 100 animals may be observed.[10]
Vocalizations
- See also: Whale song and List of whale songs
Like other whales, the male Fin Whale has been observed to make long, loud, low-frequency sounds.[11] The vocalizations of Blue and Fin Whales are the lowest known sounds made by any animal.[15] Most sounds are frequency-modulated (FM) down-swept infrasonic pulses from 16 to 40 hertz frequency (the range of sounds that most humans can hear falls between 20 hertz and 20 kilohertz). Each sound lasts between one to two seconds, and various combinations of sounds occur in patterned sequences lasting 7 to 15 minutes each. These sequences are then repeated in bouts lasting up to many days.[16] The vocal sequences have source levels of up to 184–186 decibels relative to 1 micropascal at a reference distance of one metre, and can be detected hundreds of miles from their source.[17]
When Fin Whale sounds were first recorded by US biologists, researchers did not realize that these unusually loud, long, pure, and regular sounds were being made by whales. They first investigated the possibilities that the sounds were due to equipment malfunction, geophysical phenomena, or even part of a Soviet Union scheme for detecting enemy submarines. Eventually, biologists demonstrated that the sounds were the vocalizations of Fin Whales.[15]
Direct association of these vocalizations with the reproductive season for the species and that only males make the sounds point to these vocalizations as possible reproductive displays.[18][19] Over the past 100 years, the dramatic increase in ocean noise from shipping and naval activity may have slowed the recovery of the Fin Whale population, by impeding communications between males and sexually receptive females.[20]
Habitat and migration
Summer distribution of Fin Whales in the North Pacific is the immediate offshore waters from central Baja California to Japan, and as far north as the Chukchi Sea bordering the Arctic Ocean.[23] They occur in high densities in the northern Gulf of Alaska and southeastern Bering Sea between May and October, with some movement through the Aleutian passes into and out of the Bering Sea.[24] Several whales tagged between November and January off southern California were killed in the summer off central California, Oregon, British Columbia, and in the Gulf of Alaska.[23] Fin Whales have been observed feeding in Hawaiian waters in mid-May, and several winter sightings have been made there.[25] Some researchers have suggested that the whales migrate into Hawaiian waters primarily in the autumn and winter.[26]
Although Fin Whales are certainly migratory, moving seasonally in and out of high-latitude feeding areas, the overall migration pattern is not well understood. Acoustic readings from passive-listening hydrophone arrays indicate a southward migration of the North Atlantic Fin Whale occurs in the autumn from the Labrador-Newfoundland region, south past Bermuda, and into the West Indies.[27] One or more populations of Fin Whales are thought to remain year-round in high latitudes, moving offshore, but not southward in late autumn.[27] In the Pacific, migration patterns are difficult to understand. Although some Fin Whales are apparently present in the Gulf of California year-round, there is a significant increase in their numbers in the winter and spring.[28] Antarctic Fin Whales migrate seasonally from relatively high-latitude Antarctic feeding grounds in the summer to low-latitude breeding and calving areas in the winter. The location of winter breeding areas is still unknown, since these whales tend to migrate in the open ocean and thus exact locations have been difficult to determine.[3]
Abundance and trends
North Pacific
Antarctica
Relatively little is known about the historical and current population levels of the Antarctic Fin Whale. The IWC officially estimates that the pre-whaling population of the Fin Whale in the southern hemisphere was 400,000 whales, and that the population in 1979 (at the cessation of Antarctic large scale whaling) was 85,200.[50] Both the current and historical estimates should be considered as poor estimates because the methodology and data used in the study are known to be flawed.[3] Other estimates cite current population levels of no more than 5,000 whales and possibly as low as 2,000 to 3,000.[11] As of 2006, there is no scientifically accepted estimate of current population or trends in abundance.[3]
Human interaction
- See also: Whaling and History of whaling
The Fin Whale is hunted in the Northern Hemisphere in Greenland, under the International Whaling Commission's procedure for aboriginal subsistence whaling. Meat and other products from whales killed in these hunts are widely marketed within the Greenland economy, but export is illegal. The IWC has set a quota of 19 Fin Whales per year for Greenland despite concern about uncertainty of current population levels. Iceland and Norway are not bound by the IWC's moratorium on commercial whaling because both countries filed objections to the moratorium.[3] In October 2006, Iceland's fisheries ministry authorized the hunting of nine Fin Whales through August 2007.[57] In the southern hemisphere, Japan has targeted Fin Whales in its Antarctic Special Permit whaling program for the 2005–2006 and 2006–2007 seasons at 10 whales killed per year.[58] The proposal for 2007–2008 and the subsequent 12 seasons includes 50 Fin Whales per year.[3]
Collisions with ships are an additional major cause of Fin Whale mortality. In some areas, they represent a substantial portion of the strandings of large whales. Most lethal and serious injuries are caused by large, fast-moving ships over or near the continental shelf.[59]
See also
- Whaling in Iceland
References
- ^ a b Cetacean Specialist Group (1996). Balaenoptera physalus. 2006 IUCN Red List of Threatened Species. IUCN 2006. Retrieved on 10 May 2006. Database entry includes justification for why this species is endangered
- ^ a b c d e f g h Balaenoptera physalus Fin Whale. MarineBio.org. Retrieved on 2006-10-23.
- ^ a b c d e f g h i National Marine Fisheries Service (2006). Draft recovery plan for the fin whale (<em>Balaenoptera physalus)</em> (pdf), Silver Spring, Maryland: National Marine Fisheries Service.
- ^ Revised Management Scheme. International Whaling Commission. Retrieved on 2006-11-07.
- ^ (Latin) Linnaeus, C (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.. Holmiae. (Laurentii Salvii)., 824.
- ^ Gingerich, P. (2004). "Whale Evolution", McGraw-Hill Yearbook of Science & Technology. The McGraw Hill Companies. ISBN 0071427848.
- ^ a b Bérubé, M.; A. Aguilar (1998). "A new hybrid between a blue whale, Balaenoptera musculus, and a fin whale, B. physalus: frequency and implications of hybridization". Mar. Mamm. Sci. 14: 82 – 98.
- ^ Doroshenko, V.N. (1970). "A whale with features of the fin and blue whale (in Russian)". Izvestia TINRO 70: 225–257.
- ^ Balaenoptera physalus (TSN 180527). Integrated Taxonomic Information System. Retrieved on 23 October 2006.
- ^ a b Martin, Anthony R. (1991). Whales and dolphins. London: Salamander Books.
- ^ a b c d e f g h i j k Fox, David (2001). Balaenoptera physalus (fin whale). Animal Diversity Web. Retrieved on 2006-10-22.
- ^ Tershy, B. R.; D. Wiley (1992). "Asymmetrical pigmentation in the fin whale: a test of two feeding related hypotheses". Marine Mammal Science 8 (3): 315–318.
- ^ a b c Lin, Brian (2007-06-07). Whale Has Super-sized Big Gulp. University of British Columbia. Retrieved on 2007-06-08.
- ^ Fin Whale. nature.ca: Canadian Museum of Nature. Retrieved on 2006-10-22.
- ^ a b Payne, Roger (1995). Among Whales. New York: Scribner, 176. ISBN 0-684-80210-4.
- ^ Finback Whales. Bioacoustics Research Program, Cornell Lab of Ornithology. Retrieved on 2006-10-26.
- ^ W. J. Richardson, C. R. Greene, C. I. Malme and D. H. Thomson, Marine Mammals and Noise (Academic Press, San Diego, 1995).
- ^ Croll, D.A.; Clark, C.W., Acevedo, A., Flores, S., Gedamke, J., and Urban, J. (2002). "Only male fin wales sing loud songs" (pdf). Nature 417.
- ^ Watkins, W.; Tyack, P., Moore, K., Bird, J.. "The 20 Hz signals of finback whales (Balaenoptera physalus)". The Journal of the Acoustical Society of America 82 (6): 1901–1902.
- ^ Segelken, R. (2002-06-19). Humanity's din in the oceans could be blocking whales' courtship songs and population recovery. Cornell University.
- ^ Mead, J.G. (1977). "Records of Sei and Bryde's whales from the Atlantic Coast of the United States, the Gulf of Mexico, and the Caribbean". Rep. int. Whal. Commn Spec. Iss. 1: 113–116. ISBN 0-906975-03-4.
- ^ a b Mitchell, E. (1974). "Present status of Northwest Atlantic fin and other whale stocks", in W.E. Schevill (ed.): The Whale Problem: A Status Report. Cambridge, MA: Harvard University Press, 108–169. ISBN 0-674-95075-5.
- ^ a b Rice, D.W. (1974). "Whales and whale research in the eastern North Pacific", in W.E. Schevill (ed.): The Whale Problem: A Status Report. Cambridge, MA: Harvard University Press, 170–195. ISBN 0-674-95075-5.
- ^ Reeves, R.R.; M.W. Brown (1985). "Whaling in the Bay of Fundy". Whalewatcher 19 (4): 14–18.
- ^ Mobley, J.R., Jr.; M. Smultea, T. Norris and D. Weller (1996). "Fin whale sighting north of Kaua'i, Hawai'i". Pacific Science 50 (2): 230–233.
- ^ Thompson, P.O.; W.A. Friedl (1982). "A long term study of low frequency sound from several species of whales off Oahu, Hawaii". Cetology 45: 1–19.
- ^ a b Clark, C.W. (1995). "Application of US Navy underwater hydrophone arrays for scientific research on whales". Rep. Int. Whal. Commn 45: 210–212.
- ^ Tershy, B.R.; D. Breese and C.S. Strong (1990). "Abundance, seasonal distribution and population composition of balaenopterid whales in the Canal de Ballenas, Gulf of California, Mexico". Rep. Int. Whal. Commn Spec. Iss. 12: 369–375. ISBN 0-906975-23-9.
- ^ Sigurjónsson, J. (1995). "On the life history and autecology of North Atlantic rorquals", in A.S. Blix, L. Walløe, and Ø. Ulltang (ed.): Whales, Seals, Fish and Man. Elsevier Science, 425–441. ISBN 0-444-82070-1.
- ^ D.E. Sergeant (1977). "Stocks of fin whales Balaenoptera physalus L. in the North Atlantic Ocean". Rep. Int. Whal. Commn 27: 460–473.
- ^ Allen, K.R. (1970). "A note on baleen whale stocks of the north west Atlantic". Rep. Int. Whal. Commn 20: 112–113.
- ^ Breiwick, J.M.. "Population dymanics and analyses of the fisheries for fin whales (Balaenoptera physalus) in the northwest Atlantic Ocean". (Ph.D. thesis) University of Washington, Seattle. 310 pp..
- ^ Hay, K. (1982). "Aerial line-transect estimates of abundance of humpback, fin, and long-finned pilot whales in the Newfoundland-Labrador area". Rep. Int. Whal. Commn 31: 373–387.
- ^ Kingsley, M.C.S.; R.R. Reeves (1998). "Aerial surveys of cetaceans in the Gulf of St. Lawrence in 1995 and 1996". Marine Mammal Science 17 (1): 35–75.
- ^ Larsen, F. (1995). "Abundance of minke and fin whales off West Greenland". Rep. Int. Whal. Commn 45: 365–370.
- ^ Jonsgard, A. (1974). "On whale exploitation in the eastern part of the North Atlantic Ocean", in W.E. Schevill (ed.): The Whale Problem: A Status Report. Cambridge, MA: Harvard University Press, 97–107. ISBN 0-674-95075-5.
- ^ Rørvik, C.J.; J. Sigurjónsson (1981). "A note on the catch per unit effort in the Icelandic fin whale fishery". Rep. Int. Whal. Commn 31: 379–383.
- ^ Buckland, S.T.; K.L. Cattanach and Th. Gunnlaugsson (1992). "Fin whale abundance in the North Atlantic, estimated from Icelandic and Faroese NASS-87 and NASS-89 data". Rep. Int. Whal. Commn 42: 645–651.
- ^ Rørvik, C.J.; J. Jónsson, O.A. Mathisen, and Å. Jonsgård (1976). "Fin Whales, Balaenoptera physalus (L.), Off the West Coast of Iceland. Distribution, Segregation by Length and Exploitation". Rit Fiskideildar 5: 1–30. ISSN 0484-9019.
- ^ Goujon, M.; J. Forcada and G. Desportes (1995). "Fin whale abundance in the eastern temperate North Atlantic for 1993.". Rep. Int. Whal. Commn 45: 287–290.
- ^ Buckland, S.T.; K.L. Cattanach and S. Lens (1992). "Fin whale abundance in the eastern North Atlantic, estimated from Spanish NASS-89 data". Rep. Int. Whal. Commn 42: 457–460.
- ^ Bérubé, M.; Aguilar, A., Dendanto, D., Larsen, F., Notarbartolo di Sciara, G., Sears, R., Sigurjónsson, J., Urbán-R, J. and Palsbøll, P.J. (1998). "Population genetic structure of North Atlantic, Mediterranean Sea and Sea of Cortez Fin Whales, Balaenoptera physalus (Linnaeus 1758): analysis of mitochondrial and nuclear foci". Molecular Ecology 7: 585–599. ISSN 1471-8278.
- ^ Ohsumi, S.; S. Wada (1974). "Status of whale stocks in the North Pacific, 1972". Rep. Int. Whal. Commn 24: 114–126.
- ^ Rice, D.W. (1974). "Whales and whale research in the eastern North Pacific", in W.E. Schevill (ed.): The Whale Problem: A Status Report. Cambridge, MA: Harvard University Press, 170–195. ISBN 0-674-95075-5.
- ^ Chapman, D.G. (1976). "Estimates of stocks (original, current, MSY level and MSY)(in thousands) as revised at Scientific Committee meeting 1975". Rep. Int. Whal. Commn 26: 44–47.
- ^ Barlow, J.. "Preliminary estimates of the Abundance of Cetaceans along the U.S. West Coast: 1991–2001". Administrative report LJ-03-03, available from Southwest Fisheries Science Center, 8604 La Jolla Shores Dr., La Jolla CA 92037.
- ^ Caretta, J.V., K.A. Forney, M.M. Muto, J. Barlow, J. Baker, B. Hanson, and M.S. Lowry. "U.S. Pacific Marine Mammal Stock Assessments: 2005" (PDF). U.S. Department of Commerce Technical Memorandum, NOAA-TM-NMFS-SWFSC-388.
- ^ Baretta, L.; G.L. Hunt, Jr. (1994). "Changes in the numbers of cetaceans near the Pribilof Islands, Bering Sea, between 1975–78 and 1987–89" (PDF). Arctic 47: 321–326.
- ^ Mizroch, S.A.; D.W. Rice, and J.M. Breiwick (1984). "The fin whale, Balaenoptera physalus". Mar. Fish. Review 46: 20–24.
- ^ IWC (1979). "Report of the sub-committee on protected species. Annex G, Appendix I". Rep. Int. Whal. Commn 29: 84–86.
- ^ American Cetacean Society Fact Sheet: Fin Whale, Balaenoptera physalus. American Cetacean Society. Retrieved on 2006-10-23.
- ^ a b IWC (1995). "Report of the scientific committee". Rep. Int. Whal. Commn 45: 53–221.
- ^ Barlow, J., K. A. Forney, P.S. Hill, R.L. Brownell, Jr., J.V. Caretta, D.P. DeMaster, F. Julian, M.S. Lowry, T. Ragen, and R.R. Reeves. "U.S. Pacific Marine Mammal Stock Assessments: 1996" (PDF). NOAA Technical Memo NMFD-SWFSC-248.
- ^ Yablokov, A.V. (1994). "Validity of whaling data". Nature 367: 108.
- ^ UNEP-WCMC Species Database: CITES-Listed Species. UNEP-WCMC (2006-10-23). Retrieved on 2006-10-23.
- ^ Species Profile for Finback whale. U.S. Fish & Wildlife Service. Retrieved on 2006-10-23.
- ^ Iceland to Resume Whale Hunting, Defying Global Ban. Bloomberg.com (2006-10-18). Retrieved on 2006-10-23.
- ^ U.S. Protests Japan’s Announced Return to Whaling in Antarctic. Bureau of International Information Programs, U.S. Department of State (20 November 2006). Retrieved on 2006-11-27.
- ^ Laist, D.W.; Knowlton, A.R., Mead, J.G., Collet A.S., and Podesta, M. (2001). "Collisions between ships and whales" (PDF). Marine Mammal Science 17: 35–75.
General references
- National Audubon Society Guide to Marine Mammals of the World, Reeves, Stewart, Clapham and Powell, ISBN 0-375-41141-0
- Whales & Dolphins Guide to the Biology and Behaviour of Cetaceans, Maurizio Wurtz and Nadia Repetto. ISBN 1-84037-043-2
- Encyclopedia of Marine Mammals, editors Perrin, Wursig and Thewissen, ISBN 0-12-551340-2